F3′H flavonoid 3′-hydroxylase F3′5′H flavonoid 3′5′-hydroxylase DFR dihydroflavonol 4-reductase ANS anthocyanidin synthase UFGT anthocyanidin 3-O-glucosyltransferase MYBs MYB transcription factors bHLHs basic helix-loop-helix proteins WDR WD repeat protein EBGs early biosynthetic genes LBGs late...